A preliminary study o f Cylindridia C asey , with descriptions o f new species from Middle America ( Coleóptera : Curculionidae : Baridinae )

Das bekannte Verbreitungsareal von Cylindridia Casey wird von Nord- auf Mittel- und Sudamerika ausgedehnt. Cylindridia nitidissima (Casey), C. perexilis Casey und C. simulator Casey sind neue Synonyme von C. prolixa (LeConte). Drei neue Arten werden aus hohen Lagen in Mittelamerika beschrieben: C. propinqua sp. n. aus Sudmexiko, und C. fuscipes sp. n. und C. rubripes sp. n. aus Costa Rica. Eine funfte, bislang nicht identifizierte Art wird aus Santa Catarina, Brasilien gemeldet.StichworterWeevils, new species, Cylindridia, Cyperaceae, Neotropics.Nomenklatorische Handlungenfuscipes Prena, 2006 (Cylindridia), spec. n.perexilis Casey, 1920 (Cylindridia), syn. n. of Cylindridia prolixa (LeConte, 1876)propinqua Prena, 2006 (Cylindridia), spec. n.rubripes Prena, 2006 (Cylindridia), spec. n.simulator Casey, 1920 (Cylindridia), syn. n. of Cylindridia prolixa (LeConte, 1876)nitidissima Casey, 1892 (Limnobaris), syn. n. of Cylindridia prolixa (LeConte, 1876)


Introduction
C a s e y (1892) recognized five informal subgroups (subgenera I-V) when he adopted and extended the definition of Limnobaris B e d e l for the inclusion of certain North American Baridinae.This provisional classification formed the basis for his later description of Cylindridia and five other genera (C a s e y 1920).Four species were included in Cylindridia at this occasion, and no further reference has been made to this genus ever since, except for catalogues and faunal lists.New material now extends the known distribution to the Neotropical region.The prim ary objective of this study is to contribute to the Costa Rican inventory of biodiversity.The study remains prelim inary because of its regional restriction to North and M iddle America, and the still insufficient knowledge of several width, Fig. 21-28; (4) male with deep pectoral cavity and pair of slender, anteriorly pro jecting prosternal spines usually visible from above, both structures reduced or entirely absent in depauperate specimens; (5) fore tibia in both sexes with ventrodistal spur not larger than tarsal claw; (6) claws curved and separate at base; (7) larval development in pith of flowering culms of Cyperaceae.Discussion: C ylindridia is a genus with peculiar, anteriorly projecting prosternal spines that seemed to be restricted to the Nearctic region, whereas this character state occurs rather frequently in Neotropical genera.The southward extension of the range now adds to a geographically more consistent distribution of these weevils.A n d e r s o n ) corroborates the synonymy with C. prolixa.Cylindridia similis (1 male, 3 fe males) and C. parexilis (1 female) were described from a single series collected by W ic k h a m at Greeley, Colorado ( C a s e y 1920, p. 6 4 4 ) , and lie within the variability of the material from elsewhere.The length of the male prosternai spines is variable and probably affected by the conditions imposed by the host plant and the environmental settings.
Description: Fig. 9-11, 21, 22.Total length 4.5-5.5 mm, standard length 4.2-5.2mm; derm dark green with metallic luster, glabrous, tibiae and tarsi red; rostrum as long as (male) or longer (female) than pronotum, evenly curved in both sexes, lamellar process ventrad of antennal insertion not angularly projecting; antennal club compact, approxi mately as long as distal four funicular segments combined; pronotum almost as long as wide, sides slightly rounded, constricted apically and tubulate in front; sides of aedeagus convex, apex blunt, body of aedeagus shorter than apodemes, internal sac slightly longer than apodemes, with asperities coarse in basal one-third and finer distally, basal sclerite consisting of short flagellum with pair of slender hook-like appendages, tegmen with parameres shorter than basal apodem; male sternite 9 with distal appendices very un equal.
Distribution: Mexico, Oaxaca, approximately 2150 m elevation Plant associations: Unknown Epithet: The name is a Latin adjective meaning "related".Discussion: This is the only known record for a species of Cylindridia from Mexico.W ithout dissection, the most obvious difference to the otherwise very similar Costa Rican C. rubripes is the compact antennal club and the metallic color of the femora.However, several details of the male genitalia differ significantly from those of the Costa Rican species, while agreeing much better with those of the North American C. prolixa.
C ylindridia rubripes sp.n.Description: Fig. 1, 12-15, 23, 24.Total length 2.4-5.8mm, standard length 2.3-5.6 mm; derm dark green with metallic luster, glabrous, legs red, rostrum and antenna fuscous to partially red (>2500 m elevation); rostrum approximately as long as or longer than pronotum, curved bas ally in male, nearly straight in female, lamellar process ventrad of antennal insertion not angularly projecting; antennal club elongate, at least as long as distal five funicular segments combined; sides of aedeagus subparallel, apex pointed or round (small specimens at 3000 m elevation), body of aedeagus and apodemes of sub equal length, internal sac as long as apodemes, with complex blade-like basal sclerite, tegmen with parameres slightly shorter than basal apodem; male sternite 9 with distal appendices very unequal.4-8, C. prolixar. 4-5, aedeagus, dorsal and lateral; 6, tegmen; 7-8, sternites 8 and 9. 9-11, C. propinqua-. 9-10, aedeagus, dorsal and lateral; 11, basal sclerite. 12-15, C. rubripes-. 12-14, aedeagus, dorsal and lateral (13, high-elevation  Discussion: I include under C. rubripes specimens with reddish legs and a characteristic blade-like basal sclerite in the male (Fig. 15).However, their morphological range is noteworthy.An analysis of the meristic data reveals statistically significant differences between local populations: (1) the length-width ratio of the pronotum decreases linearly with increasing altitude, (2) the standard length decreases exponentially with increas ing altitude, and (3) the length of the ante-antennal portion of the rostrum differs from site to site without obvious altitudinal effect.A gradual transition is apparent in the length-width ratio of the body of the aedeagus over the entire altitudinal range, while the apical shape changes at approximately 2500 m from triangularly pointed (Fig. 12) to round (Fig. 13); the shape is intermediate in specimens from Cuerici.At 3000 m, male specimens consistently exhibit greatly reduced prosternai spines.For distinction from the other Costa Rican species, C. fuscipes, see discussion there.

C ylindridia fu s c i p e s sp. n.
230 specimens), they can be distinguished by the color of the legs as indicated by the epithets.The outer elytral stria of C. fuscipes generally is impressed near the apex, but not so in C. rubripes.Further useful character states for separating the two species are, for fe males, the curvature of the rostrum and, for males, the presence of an angular lamellar process below the antennal insertion.Cylindridia sp.A Description: Fig. 3, 27, 28.Total length 3.5-5.6mm, standard length 3.2-5.3mm, derm reddish brown to brown, head and rostrum darker, mat, with whitish club-shaped scales, rostrum little (males) or distinctly longer (females) than pronotum, very slightly (female) or distinctly (male) curved throughout, lamellar process ventrad of antennal insertion not angularly projecting; antennal club rather compact, barely longer than distal four funicular segments combined; pronotum not quite as long as wide, sides subparallel in basal half, roundly constricted apically and subtubulate in front; sides of aedeagus paral lel, apex blunt, only middle slightly produced, apodemes shorter than body of aedeagus, internal sac short, basal sclerite absent, tegmen with parameres and basal apodem of sub equal length, male sternite 9 with distal appendices very unequal.Material examined: One series of 30 specimens from Brazil, Santa Catarina Province, Nova Teutonia, October 1974, coll.F. P l a u m a n n (CM N C) Plant associations: Unknown Fig: 1-3 : Dorsal habitus of 1, C. rubripes, male; 2, C. fu scipes, female; 3, Cylindridia sp.A, male.