Anomala felicia ARROW , 1910 and its relatives from Borneo with descriptions of six new species ( Coleoptera : Scarabaeidae : Rutelinae

The type material of Anomala felicia ARROW, 1910, Anomala kinabalensis OHAUS, 1916 and Anomala sarawakensis OHAUS, 1916 is revised. Lectotypes are selected for the nominal taxa Anomala felicia ARROW, 1910 and Anomala ovatula kinabalensis OHAUS, 1910. Anomala snizeki sp. n., Anomala kaltengensis sp. n., Anomala bifida sp. n., Anomala aequalis sp. n., Anomala mahakamensis sp. n. and Anomala manseri sp. n. are described as new to science.


Introduction
More than 60 Anomala species are described or recorded from Borneo of which the majority are Anomala species are described or recorded from Borneo of which the majority are Anomala island endemites.Only few of the Bornean Anomala species can be found also on Sumatra, Java Anomala species can be found also on Sumatra, Java Anomala or in West Malaysia.Another distribution pattern includes Borneo, Palawan and the Philippine islands (e. g. A. sulcatula BURMEISTER, 1844).Most of the Bornean BURMEISTER, 1844).Most of the Bornean BURMEISTER Anomala species were de-Anomala species were de-Anomala scribed from Sarawak and Sabah whereas material from the larger Indonesian part of the island (Kalimantan) is underrepresented in most collections.Thus, the low sampling intensity in Kalimantan does not permit final conclusions concerning the distribution pattern of the species treated in this paper (Fig. 22).OHAUS (1910) described A. kinabalensis and A. kinabalensis and A. kinabalensis A. sarawakensis, two of the species treated here, as subspecies of the Philippine Anomala ovatula OHAUS, 1910a but apart from a superficial similarity the latter is not closely related to these species.Later, Ohaus raised A. kinabalensis and A. sarawakensis to specific rank (OHAUS, 1916) and recognized their close relationship with A. felicia and provided a short identification key to these three species ( A. felicia and provided a short identification key to these three species ( A. felicia OHAUS, 1938).In the course of the examination of the unidentified Anomala material of several European museums Anomala material of several European museums Anomala and personal collecting activities in Kalimantan six new species related to the three above mentioned were discovered.In this paper these new species are described and revisions of the known taxa of the A. felicia species group are provided.

Material
The material cited in this publication is deposited in the following collections:

Diagnosis:
Anomala felicia is easily separated from the other members of this species group by the shape of Anomala felicia is easily separated from the other members of this species group by the shape of Anomala felicia the aedeagus which is simple and not elongate as in the other species.Moreover, the inner limitation of the yellow lateral patch of the pronotum is notched in the middle (Fig. 19), not evenly broad from anterior to posterior angle or extremely narrowed in the posterior half.
Distribution: Anomala felicia is only known from the Kina Balu region in the province of Sabah Anomala felicia is only known from the Kina Balu region in the province of Sabah Anomala felicia (Fig. 22).

Description and diagnosis:
Length.9.2-10.0mm; width.5.1-5.6 mm.Anomala snizeki shares most morphological characters with A. felicia as described above.The inner limitation of the yellow lateral band of the pronotum A. felicia as described above.The inner limitation of the yellow lateral band of the pronotum A. felicia is not notched but usually straight from anterior to posterior margin (Fig. 20), a character that is shared with A. kaltengensis, A. bifida, A. aequalis, A. kinabalensis and A. kinabalensis and A. kinabalensis A. mahakamensis.From these five species, A. snizeki can only be distinguished by the shape of the aedeagus.The parameres are A. snizeki can only be distinguished by the shape of the aedeagus.The parameres are A. snizeki strongly elongate whereas the ventral plate shows no extensions (Figs 3-4).

Anomala snizeki
Etymology: The species is named after Miroslav Snížek (České Budějovice) who collected some of the paratypes.Distribution: Anomala aequalis is only known from the imprecise type locality, West Borneo Anomala aequalis is only known from the imprecise type locality, West Borneo Anomala aequalis (Fig. 22).

Etymology:
The species name was chosen because A. aequalis cannot be distinguished by external characters from most of its relatives but can only be differentiated by the shape of the aedeagus.
than antennomeres 1-6 combined but longer than the funiculus.Pronotum.Ca. 1.7x as broad as long, widest at base; sides subparallel in the posterior half, strongly convergent in the anterior half; anterior angles square; posterior angles obtuse and narrowly rounded; basal bead not interrupted; punctation fine and moderately dense on disc; punctures separated by 1-2(3) diameters.Scutellum.Punctured like the disc of the pronotum.Elytra.With regular, impressed striae; intervals slightly elevated; 2nd and 3rd interstice with interrupted and not impressed secondary striae; subsutural interstice irregularly punctured anteriorly, with an indistinct stria in the posterior half.Pygidium.With sparse, rather coarse, ocellated punctation; distal margin with erect setae.Metasternum.With well-separated, ocellated punctures and rather short and sparse setosity in the lateral part.Abdominal sternites.Sparsely punctured in the middle, with denser punctures laterally; with a transverse row of setae.Protibiae.Bidentate, terminal tooth shortly bent outwards; lateral tooth small, square.Metatibiae.Rather short, a little enlarged in the middle and constricted before the apex.Claws.Modified claw of the pro-and mesotarsi apically bifid, protarsi enlarged but with angle or tooth at the inner margin.Aedeagus.Figs 1-2.