A revision of the genus Neosclerus CAMERON . Supplement I ( Coleoptera : Staphylinidae : Paederinae )

Neosclerus immutatus sp. n. (Sumatra) is described and illustrated. A recent key to the species of the genus is modified in order to account for the new species. Additional paratypes are listed for N. dupleseriatus sp. n. (N. India, central Nepal) and N. praeacutus sp. n. (Thailand). Additional records of N. fortepunctatus CAMERON, 1924 (first confirmed record) and N. granulicollis CAMERON, 1924 (new country record) from Nepal are reported. The genus now includes 27 species.


Introduction
Several months after the revision of Neosclerus had been submitted and accepted for publica-Neosclerus had been submitted and accepted for publica-Neosclerus tion, additional specimens were discovered among material of Sclerochiton KRAATZ, 1859 on loan from the Muséum d'histoire naturelle in Genève, as well as in the Franz collection at the Naturhistorisches Museum Wien.Aside from new country records of two species described a long time ago, an examination of this material yielded additional paratypes of two species described in the revision and even an undescribed species from Sumatra, from where previously no Neosclerus species had been recorded.

Material and methods
The material treated in this study is deposited in the following public institutions and private collection: The morphological studies were conducted using a Stemi SV 11 microscope (Zeiss Germany) and a Jenalab compound microscope (Carl Zeiss Jena).For the photographs a digital camera (Nikon Coolpix 995) was used.
Head length was measured from the anterior margin of the frons to the posterior mar gin of the head, elytral length at the suture from the apex of the scutellum to the posterior margin of the elytra.

Comment:
The above specimens represent the first confirmed record from Nepal.It seems possible that the females previously reported as N. brevipennis ( N. brevipennis ( N. brevipennis CAMERON, 1943) from Nepal refer to either this or the following species.Another previously unseen syntype from the original type series, but not conspecific with the lectotype, was discovered in the Franz collection at the NHMW (see the additional paratypes of N. dupleseriatus below).

Comment:
The above material shows that N. dupleseriatus is apparently not uncommon in central Nepal.It N. dupleseriatus is apparently not uncommon in central Nepal.It N. dupleseriatus also confirms that the species is wing-dimorphic.The specimen from Chakrata district belongs to the original type series of Neosclerus fortepunctatus.

Comment:
Previously, only specimens from Yunnan (China) were seen.The above material from western Thailand suggests that the species may be widespread.

Comment:
The above specimens represent the first records from Nepal.

Description:
Body length 3.2-3.5 mm.Habitus as in Fig. 1.Coloration: head, pronotum, and abdomen blackish; elytra dark-brown with the posterior margin yellow; legs yellowish; antennae reddish-yellow.Head (Fig. 2) across eyes approximately 1.2 times as wide as long; postocular region very short, almost obsolete; punctation coarse and moderately dense, slightly sparser in posterior portion of dorsal surface; interstices with distinct microreticulation and dull (Fig. 3); antenna 0.9 -1.0 mm long.Pronotum (Fig. 2) approximately 1.05 times as wide as long and 0.80-0.85times as wide as head; impunctate midline defined and rather broad; punctation as coarse and dense as that of head; interstices without microsculpture and glossy.Elytra (Fig. 2) relatively long and large, approximately 1.1 times as long as pronotum; humeral angles marked; punctation dense and fine; interstices glossy.Hind wings fully developed.Abdomen narrower than elytra; punctation fine and moderately dense on anterior tergites, somewhat less dense on posterior tergites; interstices with distinct microsculpture composed of isodiametric meshes; posterior margin of tergite VII with palisade fringe.: sternite VII with posterior margin very weakly concave, pubescence unmodified (Fig. 4); sternite VIII distinctly oblong, anteriorly unmodified and with unmodified pubescence, posterior excision acute, moderately deep, and V-shaped, its depth approximately 1/6 the length of sternite (Fig. 5); aedeagus small, approximately 0.45 mm long, with dorso-ventrally flattened ventral process; internal sac with two long series of small sclerotized structures (Figs 6-7).species, N. immutatus is readily separated by the much darker coloration of the body, the much N. immutatus is readily separated by the much darker coloration of the body, the much N. immutatus less dense punctation of the head and pronotum, the more transverse head, and by the completely different shape of the ventral process of the aedeagus.In order to account for the new species, the key in ASSING ( 2011

Distribution and bionomics:
The type locality is situated at the eastern shore of lake Toba a few kilometres to the south of Prapat in the north of Sumatra.The specimens were collected in February, either from under bark of a dead tree or from between stones and gravel in a dry river bed (SCHILLHAMMER, pers.SCHILLHAMMER, pers.SCHILLHAMMER comm.).