Breddin ' s types of Ochlerus ( Hemiptera , Pentatomidae , Discocephalinae )

After more than a century, the type specimens of the Pentatomidae genus Ochlerus Spin o la , 1837 described by Gustav Breddin are revised. Due to unfortunate circumstances, the author left most of the related work incomplete and it resulted in the possible mislabeling of the type material, as well as a change in the institutions where they should be deposited. Types are assigned to the following species: Ochlerus bergrothi, O. bistillatus, O. communis, O. cotylophorus, O. dentijugis, O. handlirschi, O. incisulus, O. notatulus, O. profanus, O. rusticus, O. signoreti, O. stylulatus, and O. tenuicornis. The male of O. tenuicornis is transferred to a new species, Ochlerus breddini.


Introduction
With a worldwide distribution (except in polar regions), Pentatomidae is listed as the fourth in diversity among the Heteroptera (G r a zia et al. 2012). These insects are highly related to cultivated plants such as soybeans and palm trees, among others, and constantly reported as plagues, which highlight its impact to economy (Pa n iz z i et al. 2000). The family is divided into 10 subfamilies, six of which occur in the Neotropical region (Rid e r , 2014).
One of such subfamilies is Discocephalinae, where the genus we deal with in the present paper is placed. Ochlerus Spin o l a , 1837 has a record of unresolved taxonomy and classification matters, having its monophyly recently disputed (Cam po s & G r a z ia , 2006). Being the type genus for the tribe Ochlerini, surprisingly it has not received significant attention from taxonomists and systematists alike, besides the diagnosis by Rolston Tab. 1 : List of valid species of Ochlerus in alphabetical order.

Ochlerus cinctus Spinola 1837
Ochlerus circummaculatus Stâl 1860 Ochlerus communis Breddin 1910 Ochlerus coriaceus Herrich-Schäffer 1844 Ochlerus cotylophorus Breddin 1910 Ochlerus dentijugis Breddin 1910 Ochlerus handlirschi Breddin 1910 Ochlerus incisulus Breddin 1910 Ochlerus lutosus Herrich-Schäffer 1844 Ochlerus notatulus Breddin 1910 Ochlerus profanus Breddin 1910 Ochlerus rusticus Breddin 1910 Ochlerus signoreti Breddin 1910 Ochlerus stylulatus Breddin 1910 Ochlerus tenuicornis Breddin 1910Breddin (1992 Type locality: Unknown. Comments: Breddin (1910) based his description on five males, deposited at the NMW (4 specimens) and the Hamburg Museum (one specimen). In addition, he associated one female deposited at the NMW, but he indi cated the identification as doubtful. We were able to find one male deposited at the NMW. There is a fourth specimen from Peru which was found next to these other three and may be one of the syntypes. It has a matching general morphol ogy, despite the fact that it lacks the abdomen; since the latter is a diagnostic structure we decided to not include it as a lectotype or syntype.

Comments:
The diagnostic feature of the lateral projec tion on the mandibular plates and the shape of the pygophore, much different from the other species, lead us to the conclusion that this specimen is a syntype of O. dentijugis. Breddin stated that the specimen on which he based his description originated from Colombia, however the label pinned with the syntype states other wise, again leading to the possibility that some labels are mixed and/or misplaced.
3.1.6. Ochlerus handllrschl Breddin, 1910 (pp.  Etimology: The name of the species is an homage to the late Gustav Breddin, a researcher who made significant contributions to Ochlerus and who described most of its currently known species.
Diagnosis: Body light to dark brown, ventral facies yellow speckled. Rostrum no surpassing urosternite V. Antero lateral angles of the pronotum slightly produced laterad with a rounded apex. Small yellow maculae laterad to apex of radial veins. For a more accurate diagnosis, see description of genitalia.

Final Considerations
Despite the uncertainty to respective whereabouts, most of the missing specimens may still be found in the aforementioned collections should a hemipterologist examine them. We also conclude that the death of Breddin left a few gaps in the type specimen organization, but in the big picture there were not as many mistakes as we could expect from an incomplete labeling process.
The decision on where the misdeposited specimens should be sent will be left for the respective curators and may be addressed in the near future. Finally, we hope that this study can help in the identification of speci mens of Ochlerus, as well as in other studies delving into Breddin's posthmous works.