Nilsson, A.N., Kholin, S.K. and Minakawa, N. (1999) “The diving beetles of Kamchatka, with additional records from Sakhalin and the Kuril Islands (Coleoptera: Dytiscidae)”., Contributions to Entomology = Beiträge zur Entomologie, 49(1), pp. 107–131. doi:10.21248/contrib.entomol.49.1.107-131.