B. Seifert, “Lasius nigroemarginatus Forel, 1874 is a F1 Hybrid between L. emarginatus (Olivier, 1792) and L. platythorax Seifert, 1991 (Hymenoptera, Formicidae)”, contrib.entomol., vol. 69, no. 2, pp. 291–300, Dec. 2019, doi: 10.21248/contrib.entomol.69.2.291-300.